The World's 25 Most Endangered Primates
The World's 25 Most Endangered Primates is a list of highly endangered primate species selected and published by the International Union for Conservation of Nature Species Survival Commission Primate Specialist Group (IUCN/SSC PSG), the International Primatological Society (IPS), and Conservation International (CI). The IUCN/SSC PSG worked together with CI to start the list in 2000, but in 2002, during the 19th Congress of the International Primatological Society, primatologists reviewed and debated the list, resulting in the 2002–2004 revision and the endorsement of the IPS. The publication has since been a joint project between the three conservation organizations and has been revised every two years following the biannual Congress of the IPS.[1] Starting with the 2004–2006 report, the title changed to Primates in Peril: The World's 25 Most Endangered Primates.[2] That same year, the list began to provide information about each species, including their conservation status and the threats they face in the wild.[1] The species text is written in collaboration with experts from the field, with 60 people contributing to the 2006–2008 report[3] and 85 people contributing to the 2008–2010 report.[1] The 2004–2006 and 2006–2008 reports were published in the IUCN/SSC PSG journal Primate Conservation,[2][4] while the 2008–2010 report was published as an independent publication by all three contributing organizations.[1]
The 25 species on the 2008–2010 list are distributed between 17 countries. The countries with the most species on the list are Madagascar (five species), Vietnam (five species), and Indonesia (four species). The list is broken into four distinct regions: the island of Madagascar, the continent of Africa, the continent of Asia including the islands of Indonesia, and the Neotropics (Central and South America). Seven species have been on all five published lists: the silky sifaka (Propithecus candidus), Delacour's langur (Trachypithecus delacouri), golden-headed langur (Trachypithecus poliocephalus poliocephalus), grey-shanked douc (Pygathrix cinerea), Tonkin snub-nosed monkey (Rhinopithecus avunculus), Cross River gorilla (Gorilla gorilla diehli), and Sumatran orangutan (Pongo abelii).[1]
The purpose of the list, according to Russell Mittermeier, the president of CI, is "to highlight those [primate species] that are most at risk, to attract the attention of the public, to stimulate national governments to do more, and especially to find the resources to implement desperately-needed conservation measures."[5] Species are selected for the list based on two primary reasons: extremely small population sizes and very rapid drops in numbers. These reasons are heavily influenced by habitat loss and hunting, the two greatest threats that primates face. More specifically, threats listed in the report include deforestation due to slash and burn agriculture, clearing for pasture or farmland, charcoal production, firewood production, illegal logging, selective logging, mining, land development, and cash crop production; forest fragmentation; small population sizes; live capture for the exotic pet trade; and hunting for bushmeat and traditional medicine.[1]
Key
Species |
Common and scientific name of the species, including a picture if available |
Years listed |
Years the species has been included in the IUCN's list of the "Top 25 Most Endangered Primates" |
Location(s) |
Countries in which it is found |
Estimated population |
Latest population estimate from the IUCN |
IUCN status |
Conservation status of the species, per the IUCN as of the date of the latest list publication |
Threats |
A list of threats facing the species; used by the IUCN in assessing conservation status |
Current list
The World's 25 Most Endangered Primates, 2008–2010[1]
Species |
Years listed |
Location(s) |
Estimated population |
IUCN status |
Threats |
Madagascar |
Greater bamboo lemur
Prolemur simus |
2002
2004
2006
2008 |
Madagascar |
100–160 or fewer |
1 !
Critically Endangered[6] |
- small, isolated populations
- loss of habitat and fragmentation (slash-and-burn agriculture, mining, illegal logging, cutting of bamboo)
- hunting (bushmeat)
- reduced availability of drinking water due to climatic change
- extreme dietary specialization and dependency on giant bamboo
|
Gray-headed lemur
Eulemur cinereiceps |
2004
2006
2008 |
Madagascar |
7,265 ± 2,268 |
1 !
Endangered[7] |
- very small range (~700 km2)
- hybridization with Red-fronted Lemur (E. rufifrons)
- low population densities
- habitat loss and fragmentation (fragmented, small populations)
- cyclones
- hunting (bushmeat)
|
Blue-eyed black lemur
Eulemur flavifrons |
2008 |
Madagascar |
450–2,300 |
1 !
Endangered[8] |
- very small range (~2,700 km2)
- loss of habitat (slash-and-burn agriculture, selective logging)
- hunting (bushmeat)
- live capture (pet trade)
|
Northern sportive lemur
Lepilemur septentrionalis |
2008 |
Madagascar |
fewer than 100 |
1 !
Critically Endangered[9] |
- very small range
- habitat loss (charcoal production)
- hunting (bushmeat)
|
Silky sifaka
Propithecus candidus |
2000
2002
2004
2006
2008 |
Madagascar |
100–1,000 |
1 !
Critically Endangered[10] |
- very small range
- hunting (bushmeat)
- habitat loss (slash-and-burn agriculture, selective logging, firewood)
|
Africa |
Rondo dwarf galago
Galagoides rondoensis |
2006
2008 |
Tanzania |
unknown |
1 !
Critically Endangered[11] |
- very small range
- habitat loss and fragmentation (agricultural encroachment, charcoal production, logging)
|
Roloway monkey
Cercopithecus diana roloway |
2002
2006
2008 |
Côte d'Ivoire
Ghana |
unknown |
1 !
Endangered[12] |
- hunting (bushmeat)
- habitat loss and fragmentation (numerous documented local extinctions)
|
Tana River red colobus
Procolobus rufomitratus |
2002
2004
2006
2008 |
Kenya |
fewer than 1,000 |
1 !
Endangered[13] |
- very small range
- habitat loss and fragmentation (agricultural encroachment, selective logging for local use [houses, canoes])
- exploitation of non-timber products
- parasitic infection of isolated populations
|
Niger Delta red colobus
Procolobus epieni |
2008 |
Nigeria |
unknown |
1 !
Critically Endangered[14] |
- very small range (~1,500 km2)
- hunting (bushmeat)
- habitat loss and degradation (logging of important food trees, loss of marsh forests due to canal construction)
|
Kipunji
Rungwecebus kipunji |
2006
2008 |
Tanzania |
around 1,117 |
1 !
Critically Endangered[15] |
- very small range
- habitat loss and fragmentation
- hunting (bushmeat)
|
Cross River gorilla
Gorilla gorilla diehli |
2000
2002
2004
2006
2008 |
Cameroon
Nigeria |
200–300 |
1 !
Critically Endangered[16] |
- small, restricted range
- habitat loss (agricultural encroachment, fires to clear forest or improve pasture, development activities [roads])
- hunting (bushmeat, wire snares set for other wildlife)
|
Asia |
Siau Island tarsier
Tarsius tumpara |
2006
2008 |
Indonesia (Siau Island) |
Low thousands at best |
Not Evaluated |
- island population (near an active volcano)
- very small range
- high human density
- hunting (bushmeat [used as snack food])
- habitat degradation
|
Javan slow loris
Nycticebus javanicus |
2008 |
Indonesia (Java) |
unknown |
1 !
Endangered[17] |
- live capture (pet trade [intense])
- hunting (traditional medicine [intense])
- habitat loss (agriculture, development activities [roads], human disturbance)
|
Pig-tailed langur
Simias concolor |
2002
2004
2006
2008 |
Indonesia (Mentawai Islands) |
around 3,347 |
1 !
Critically Endangered[18] |
- island population
- habitat loss (human encroachment, product extraction, commercial logging, conversion to cash crops and oil palm plantations)
- hunting (bushmeat)
|
Delacour's langur
Trachypithecus delacouri |
2000
2002
2004
2006
2008 |
Vietnam |
200 or fewer |
1 !
Critically Endangered[19] |
- restricted range (400–450 km2)
- habitat fragmentation (60% occur in isolated populations of fewer than 20 animals)
- hunting (traditional medicine [for trade in bones, organs and tissues])
|
Golden-headed langur
Trachypithecus poliocephalus poliocephalus |
2000
2002
2004
2006
2008 |
Vietnam |
60–70 |
1 !
Critically Endangered[20] |
- island population (karst island of 140 km2)
- habitat fragmentation (seven isolated subpopulations)
- hunting (traditional medicine [for trade in bones, organs and tissues])
|
Western purple-faced langur
Trachypithecus vetulus nestor |
2004
2006
2008 |
Sri Lanka |
unknown |
1 !
Critically Endangered[21] |
- habitat loss and fragmentation (90% of forest in its range has been lost to urbanization and agriculture)
- dependent on gardens for survival
- live capture (pet trade)
- hunting (persecution as pests)
- other human factors (electrocution [power lines], road kill, dog attacks)
|
Grey-shanked douc
Pygathrix cinerea |
2000
2002
2004
2006
2008 |
Vietnam |
600–700 |
1 !
Critically Endangered[22] |
- restricted range
- habitat loss and fragmentation (agriculture, logging, fuelwood)
- hunting (bushmeat)
|
Tonkin snub-nosed monkey
Rhinopithecus avunculus |
2000
2002
2004
2006
2008 |
Vietnam |
fewer than 200 |
1 !
Critically Endangered[23] |
- restricted range (five isolated localities)
- habitat loss and fragmentation (logging, shifting cultivation, dam construction, influx of human populations)
- hunting (bushmeat [increasing])
|
Eastern black crested gibbon
Nomascus nasutus |
2008 |
China
Vietnam |
around 110 |
1 !
Critically Endangered[24] |
- small, isolated populations
- very small range (~48 km2)
- habitat loss, fragmentation, and disturbance (cultivation, pasture, firewood, charcoal production)
|
Western hoolock gibbon
Hoolock hoolock |
2006
2008 |
Bangladesh
India
Myanmar |
fewer than 5,000 |
1 !
Endangered[25] |
- very small populations (recent, very rapid declines in numbers)
- habitat loss and fragmentation (human encroachment, tea plantations, slash-and-burn cultivation)
- hunting (bushmeat, traditional medicine)
- live capture (pet trade)
|
Sumatran orangutan
Pongo abelii |
2000
2002
2004
2006
2008 |
Indonesia (Sumatra) |
around 6,600 |
1 !
Critically Endangered[26] |
- recent, very rapid declines in numbers
- only 10 fragmented habitat units
- habitat loss and fragmentation (fires, agriculture and oil palm plantations, roads, logging, encroachment)
- hunting (pests, bushmeat) [occasional]
- live capture (pet trade) [occasional]
|
Neotropics |
Cotton-top tamarin
Saguinus oedipus |
2008 |
Colombia |
fewer than 6,000 |
1 !
Critically Endangered[27] |
- habitat loss and fragmentation (large-scale agricultural production [cattle] and farming, logging, oil palm plantations, hydroelectric projects)
- live capture (pet trade [current], biomedical research [past])
|
Variegated spider monkey
Ateles hybridus |
2004[N 1]
2006
2008 |
Colombia
Venezuela |
unknown |
1 !
Critically Endangered[28] |
- restricted ranges of two subspecies
- low population densities
- habitat loss and fragmentation (agriculture, cattle-ranching)
- hunting (bushmeat)
- live capture (pet trade)
|
Yellow-tailed woolly monkey
Oreonax flavicauda |
2000
2006
2008 |
Peru |
unknown |
1 !
Critically Endangered[29] |
- restricted range
- low population densities
- habitat loss (agriculture, logging, roads, colonization)
- hunting (bushmeat, fur)
- live capture (pet trade)
|
Former list members
With each new publication, species are both added and removed from the list. In some cases, removal from the list signifies improvement for the species. With the publication of the 2006–2008, four species were removed from the list because of increased conservation efforts: the black lion tamarin (Leontopithecus chrysopygus), golden lion tamarin (Leontopithecus rosalia), mountain gorilla (Gorilla beringei beringei), and Perrier's sifaka (Propithecus perrieri).[30] In 2008, the black lion tamarin went from Critically Endangered to Endangered and the golden lion tamarin was similarly promoted in 2003 after three decades of collaborative conservation efforts by zoos and other institutions. Well-protected species such as these still have very small populations, and due to deforestation, new habitat is still needed for their long-term survival.[5] The Hainan black crested gibbon (Nomascus hainanus), which was removed from the 2008–2010 list, still has fewer than 20 individuals left, but significant efforts to protect it are now being made.[1] Mittermeier claimed in 2007 that all 25 species would be elevated off the list within five to ten years if conservation organizations had the necessary resources.[30]
Unlike the changes in the 2006–2008 report, not all species were removed from the 2008–2010 list due to improvement in their situation. Instead, new species were added to bring attention to other closely related species with very small populations that are also at risk of extinction. For example, the highly endangered eastern black crested gibbon (Nomascus nasutus) replaced the Hainan black crested gibbon. The Javan slow loris (Nycticebus javanicus) replaced the Horton Plains slender loris (Loris tardigradus nycticeboides) because the former has been hit the hardest of Asian lorises, all of which are declining rapidly due primarily to capture for the exotic pet trade, as well as use in traditional medicines and forest loss. In another case, the brown-headed spider monkey (Ateles fusciceps fusciceps) was omitted from the list since no spokesperson could be found for the species.[1]
Primates formerly listed in the "Top 25 Most Endangered Primates"[1]
Species |
Years listed |
Location(s) |
Estimated population |
IUCN status |
Threats |
Madagascar |
Golden-crowned sifaka
Propithecus tattersalli |
2000 |
Madagascar |
6,000–10,000[31] |
1 !
Endangered[31] |
- hunting (by gold miners)
- loss of habitat (slash-and-burn agriculture, uncontrolled grass fires, wood extraction [housing & firewood], selective logging, gold mining)[31]
|
Golden bamboo lemur
Hapalemur aureus |
2000 |
Madagascar |
fewer than 5,916[32] |
1 !
Endangered[32] |
- loss of habitat (slash-and-burn agriculture, cutting of bamboo [for building houses, carrying water, making baskets and other local uses])
- hunting (bushmeat)[32]
|
Lac Alaotra bamboo lemur
Hapalemur alaotrensis |
2000 |
Madagascar |
around 2,500[33] |
1 !
Critically Endangered[33] |
- loss of habitat (agricultural encroachment, burning of marshlands [to catch fish and for cattle grazing]
- hunting (bushmeat)
- live capture (local pet trade)[33]
|
Sahamalaza sportive lemur
Lepilemur sahamalazensis |
2006 |
Madagascar |
unknown |
1 !
Data Deficient[34] |
- loss of habitat (agricultural encroachment, charcoal production, selective logging for local use [houses])
- hunting (bushmeat)[34]
|
Perrier's sifaka
Propithecus perrieri |
2000
2002
2004 |
Madagascar |
around 915[35] |
1 !
Critically Endangered[35] |
- loss of habitat (slash-and-burn agriculture, charcoal production, fires to clear forest for pasture, mining)
- hunting (bushmeat)[35]
|
Africa |
Mt. Rungwe galago
Galagoides sp. |
2004 |
Tanzania |
unknown |
Not Evaluated |
- loss of habitat (logging, agricultural encroachment, charcoal production)
- hunting (bushmeat)[2]
|
Sclater's guenon
Cercopithecus sclateri |
2000 |
Nigeria |
unknown |
1 !
Vulnerable[36] |
- habitat loss and fragmentation (logging, agricultural encroachment, oil exploration)
- high human density
- hunting (bushmeat)[36]
|
Drill
Mandrillus leucophaeus |
2000 |
Cameroon
Equatorial Guinea (Bioko)
Nigeria |
unknown |
1 !
Endangered[37] |
- small range
- loss of habitat (clearcutting [for chipboard factories and settlement])
- hunting (bushmeat, persecution as pests)[37]
|
Tana River mangabey
Cercocebus galeritus galeritus |
2002 |
Kenya |
1,000–1,200[38] |
1 !
Endangered[38] |
- loss of habitat (palm oil production, logging, agricultural encroachment, grass fires intended to prevent forest regeneration, overgrazing, damming and irrigation projects)
- hunting (persecution as pests)[38]
|
Sanje mangabey
Cercocebus sanjei |
2000
2002
2004 |
Tanzania |
fewer than 1,300[39] |
1 !
Endangered[39] |
- loss of habitat (logging, charcoal production)
- hunting (persecution as pests)[39]
|
Sooty mangabey
Cercocebus atys lunulatus |
2000
2002
2004 |
Côte d'Ivoire
Ghana |
unknown |
1 !
Endangered[40] |
- habitat loss and degradation
- hunting (bushmeat)[40]
|
Miss Waldron's red colobus
Piliocolobus badius waldronae |
2000
2002
2006 |
Côte d'Ivoire
Ghana |
unknown |
1 !
Critically Endangered[41] |
- very small populations (recent, very rapid declines in numbers)
- habitat loss
- hunting (bushmeat)[41]
|
Pennant's colobus
Procolobus pennantii |
2004
2006 |
Congo
Equatorial Guinea (Bioko)
Nigeria |
unknown
(P. p. bouvieri may be extinct)[42] |
1 !
Critically Endangered[42] |
- habitat degradation
- hunting (bushmeat)[42]
|
Mountain gorilla
Gorilla beringei beringei |
2000
2002
2004 |
Rwanda
Uganda |
around 600 |
1 !
Critically Endangered[43] |
- two isolated populations
- political instability
- human diseases
- hunting (bushmeat)[43]
|
Asia |
Horton Plains slender loris
Loris tardigradus nycticeboides |
2004
2006 |
Sri Lanka |
unknown |
1 !
Endangered[44] |
- five isolated populations
- habitat loss
- hunting (bushmeat)[44]
|
Natuna Island surili
Presbytis natunae |
2002 |
Indonesia |
fewer than 10,000[45] |
1 !
Vulnerable[45] |
- two isolated populations
- habitat loss and degradation
- live capture (pet trade)[45]
|
White-headed langur
Trachypithecus poliocephalus leucocephalus |
2002 |
China |
unknown |
1 !
Critically Endangered[46] |
- very small populations (recent, very rapid declines in numbers)
- habitat loss
- hunting[46]
|
Miller's grizzled langur
Presbytis hosei canicrus |
2004 |
Indonesia (Kalimantan) |
unknown |
1 !
Endangered[47] |
- habitat loss and fragmentation
- hunting[47]
|
Black snub-nosed monkey
Rhinopithecus bieti |
2002 |
China |
fewer than 2,000[48] |
1 !
Endangered[48] |
- habitat loss (logging, fires for agricultural use, pasture)
- pesticide use
- hunting (non-targeted [snares])[48]
|
Gray snub-nosed monkey
Rhinopithecus brelichi |
2002 |
China |
around 750[49] |
1 !
Endangered[49] |
- one isolated population (vulnerable to epidemic disease or catastrophes)
- habitat loss (forest clearing, development for tourism, agricultural expansion, firewood)
- hunting (non-targeted)[49]
|
Silvery gibbon
Hylobates moloch |
2000 |
Indonesia (Java) |
4,000–4,500[50] |
1 !
Endangered[50] |
- habitat loss and fragmentation
- live capture (pet trade)[50]
|
Hainan black crested gibbon
Nomascus hainanus |
2000
2004
2006 |
China (Hainan) |
around 20[51] |
1 !
Critically Endangered[51] |
- extremely small population size
- suboptimal, protected habitat
- possible gender bias in recent births
- hunting (bushmeat)[51]
|
Neotropics |
Golden lion tamarin
Leontopithecus rosalia |
2000 |
Brazil (Rio de Janeiro) |
more than 1,000[52] |
1 !
Endangered[52] |
- habitat loss and fragmentation (fires to clear forest for pasture)
- live capture (pet trade)[52]
|
Black lion tamarin
Leontopithecus chrysopygus |
2000 |
Brazil (São Paulo) |
around 1,000[53] |
1 !
Endangered[53] |
- small population size (11 isolated populations, but only one is viable)
- habitat loss and fragmentation[53]
|
Superagui lion tamarin
Leontopithecus caissara |
2000
2002
2004 |
Brazil (Paraná and São Paulo) |
fewer than 400[54] |
1 !
Critically Endangered[54] |
- small, isolated populations
- habitat loss and degradation (agricultural encroachment, palm heart harvesting, tourism)
- high human density (increased squatting by impoverished people, land speculation)
- hunting (bushmeat)[54]
|
Golden-bellied capuchin
Cebus xanthosternos |
2000
2002
2004 |
Brazil (Bahia, Minas Gerais?) |
unknown |
1 !
Critically Endangered[55] |
- habitat loss
- hunting (bushmeat)[55]
|
Brown-headed spider monkey
Ateles fusciceps fusciceps |
2006 |
Colombia (mainland)
Ecuador (mainland)
Panama |
unknown |
1 !
Critically Endangered[56] |
- very small populations (80% declines in numbers over 45 years)
- habitat loss
- hunting (bushmeat)[56]
|
Northern muriqui
Brachyteles hypoxanthus |
2000
2002
2004 |
Brazil (Bahia, Espírito Santo, Minas Gerais) |
more than 855[57] |
1 !
Critically Endangered[57] |
- small, isolated populations
- habitat loss and fragmentation
- hunting (bushmeat [past], sport [past])[57]
|
List history
With the exception of the 2000–2002 publication, which was written collaboratively by the IUCN/SSC PSG and CI, the list has been revised every two years following the biannual Congress of the IPS. The 2002–2004 list resulted from the 19th Congress of the IPS in Beijing, China; the 2004–2006 list followed the 20th Congress of the IPS, held in Torino, Italy; the 2006–2008 list after the 21st Congress in Entebbe, Uganda; and the 2008–2010 list followed the 22nd Congress held in Edinburgh, UK.[1]
The 2008 IUCN Red List of Threatened Species offered assessments of 634 primate taxa, of which 303 (47.8%) were listed as threatened (Vulnerable, Endangered, or Critically Endangered). A total of 206 primate species were ranked as either Critically Endangered or Endangered, 54 (26%) of which have been included at least once in The World's 25 Most Endangered Primates since 2000.[1]
Notes
- ^ a b c The subspecies Ateles hybridus brunneus was listed in the 2004–2006 report, but the listing was expanded to cover both subspecies of Ateles hybridus starting with the 2006–2008 report.[1][2][4]
- ^ In the 2000–2002 report, this lemur was listed as Propithecus diadema candidus, but it has since been reclassified as a separate species, Propithecus candidus.[1][58][59]
- ^ In the 2000–2002 report, this lemur was listed as Propithecus diadema perrieri, but it has since been reclassified as a separate species, Propithecus perrieri.[1][58][59]
- ^ In the 2000–2002 report, this lemur was listed as Hapalemur griseus alaotrensis, but it has since been reclassified as a separate species, Hapalemur alaotrensis.[1][58]
- ^ a b In the 2000–2002 and 2004–2006 reports, this gorilla was listed as Gorilla beringei, but other reports instead used Gorilla b. beringei.[1][2][58][59]
- ^ In the 2000–2002 and 2002–2004 reports, the mangabey was listed as Cercocebus galeritus sanjei, but it has since been classified as a separate species, Cercocebus sanjei.[1]
- ^ a b c In the 2000–2002, 2002–2004, and 2006–2008 reports, this colobus monkey was listed as Procolobus badius waldroni, but the spelling of the name has since been corrected to waldronae.[58][59][60]
- ^ a b In the 2000–2002 and 2002–2004 reports, this monkey was listed as Trachypithecus poliocephalus, but with other subspecies now recognized by the IUCN, it is now known as Trachypithecus p. poliocephalus.[1][2][58][59]
- ^ a b In the 2000–2002 and 2002–2004 reports, this douc was listed as Pygathrix nemaeus cinerea, but has since been recognized as a separate species, Pygathrix cinerea.[1][2]
- ^ a b In the 2000–2002 and 2004–2006 reports, this gibbon was listed as Hylobates concolor hainanus, but it has since been recognized as a separate species and placed in the genus Nomascus, so that it is known as Nomascus hainanus.[1][58]
- ^ In the 2000–2002 report, this species was listed as Lagothrix flavicauda, but it has since been renamed to Oreonax flavicauda.[1]
- ^ In the 2002–2004 report, this monkey was listed as Trachypithecus leucocephalus, but it is now considered only a subspecies, Trachypithecus poliocephalus leucocephalus.[1][59]
- ^ a b In the 2004–2006 and 2006–2008 reports, this species was listed as Eulemur albocollaris, but it has since been renamed to Eulemur cinereiceps.[1]
- ^ In the 2006–2008 report, this tarsier was listed as "Tarsius sp.", but it has since been formally named as Tarsius tumpara.[1][4]
- ^ The species Ateles fusciceps was listed in the 2006–2008 report, but the common name and description match the later identified subspecies, Ateles fusciceps fusciceps.[4]
References
- ^ a b c d e f g h i j k l m n o p q r s t u v w x y z Mittermeier, R.A.; Wallis, J.; Rylands, A.B. et al., eds (2009) (PDF). Primates in Peril: The World's 25 Most Endangered Primates 2008–2010. Illustrated by S.D. Nash. IUCN/SSC Primate Specialist Group (PSG), International Primatological Society (IPS), and Conservation International (CI). pp. 1–92. ISBN 978-1-934151-34-1. http://www.primate-sg.org/PDF/Primates.in.Peril.2008-2010.pdf.
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External links
The World's 25 Most Endangered Primates
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